TC question 1

Ethologists are convinced that many animals survive through learning - but learning that is ----- their genetic programming , learning as thoroughly ---- as the most instinctive of behavioral responses.

Can u help me to find the clue and answer choices with explaination

I believe the clue answer is “deep rooted, ingrained, entrenched” coming from the word genetically and instinctually which means it is not learned behavior.
Do any options match these synonyms?